Prime Minister Boris Johnson and his fiancee Carrie Symonds have announced the birth of a son, a spokeswoman for the PM and his partner said both mother and baby are "doing very well".
It is understood the PM, who has just recovered from coronavirus, was present throughout the birth, at an NHS hospital in London.
"The PM and Ms Symonds would like to thank the fantastic NHS maternity team," the PM's spokeswoman added.
The couple has received messages of congratulation from across the political spectrum, and the PM's father Stanley said he was "absolutely delighted" and "thrilled" by the birth of his grandson.
They are the first unmarried couple to move into Downing Street together.
The baby is Ms Symonds' first child, while Mr Johnson is known to have fathered five.
Mr Johnson returned to work on Monday, after a battle with coronavirus which saw him spend three nights in intensive care. Ms Symonds also suffered symptoms of the disease.
Ms Symonds said on social media that she had spent a "worrying" week in bed with the symptoms of the virus while the PM was self-isolating with the disease.
She later sent Mr Johnson baby scans and daily messages while he was in the hospital, to keep his morale up.
Mr Johnson's Conservative colleagues have also been congratulating the couple on social media, with Health Secretary Matt Hancock saying: "So thrilled for Boris and Carrie. Wonderful to have a moment of unalloyed joy!"
Labour leader Sir Keir Starmer also offered his congratulations on the "wonderful news".
Scottish First Minister Nicola Sturgeon tweeted: "Some good news - sending congratulations to Carrie and the PM. And wishing health and happiness to the wee one.
Arlene Foster, the First Minister of Northern Ireland, congratulated the couple, adding: "More sleepless nights ahead!" Mr Johnson has previously indicated he will take paternity leave but it is not known if his plans have changed, in light of the coronavirus emergency.
The new arrival is the third baby born to a serving prime minister in recent history.
Tony Blair's wife Cherie gave birth to son Leo in May 2000, three years after her husband's first election victory, and David Cameron and wife Samantha welcomed daughter Florence in 2010.
Commons Speaker Sir Lindsay Hoyle congratulated Mr Johnson and Ms Symonds "on behalf of everyone in the House of Commons," adding: "Such happy news amid so much uncertainty - 2020 is certainly a year they will never forget."
The Archbishop of Canterbury, Justin Welby, tweeted his congratulations and prayers for the couple. "as they welcome their son into the world Wishing them every blessing and happiness."
There was speculation that Mr Johnson would take part in his first Prime Minister's Questions since recovering from coronavirus later on Wednesday.
But his place will be taken Foreign Secretary Dominic Raab, who has been deputising for him.
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